I've inherited a model that runs without errors when opened and executed in edit mode. However, when I simply open the model and run it directly, it throws an error. Since it works in edit mode, the usual debugging method of observing what turns red isn't helpful. I'm struggling to pinpoint the source of the error.

Is there a way to modify the model, to help me debug the model, and track its flow and narrow down the location of the error?
The model itself is huge, and I would rather not have to rebuild it. Based on the limited messages, it appears to be failing in the later third of the model.

Any help on how to debug the model would be greatly appreciated.