In simple terms, the answer for the original question is this:
The user controls the minimum drainage area required to form the beginning of a reach. This is the same as defining the average areas of the headwater sub-basins. This is done when you set the threshold in the stream definition.
After that, the user does not have control over the size of the downstream sub-basins, because the distance between reaches, or streams, is what drives it. Hence, the downstream sub-basins can be much larger, or much smaller than the initial threshold set on the stream definition.
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